Trochamminoides grzybowskii, nom.nov., a new name for Trochammina elegans Grzybowski, 1898 (Foraminiferida)

INTRODUCTION Trochamminoides elegans (Grzybowski) [= Trochammina elegans Grzybowski, 1898] is a familiar species that is commonly recorded from the Paleogene of the Carpathian flysch. However, during the course of work on a taxonomic revision of the Grzybowski Collection, it has become apparent that the name of this taxon is in actuality, invalid. The designation Trochammina elegans was first used by Rzehak (1887), but since the name only appeared in a table (without a description or illustration) the name is unavailable under Article 12 of the ICZN. Grzybowski (1898), who had received specimens from Rzehak, referred to this species as “Trochammina elegans Rzehak”, but the authorship of the species belongs to Grzybowski under Article 50 of the ICZN. Grzybowski (1898) described Trochammina elegans as follows: Test agglutinated, surface rough; circular in outline, spirally coiled in two and one-half to three whorls. Chambers spherical, increasing in size, numerous, with depressed sutures; there are thirteen to fifteen of them in the last whorl. Aperture at the lower edge of the last chamber, just above the rounded periphery. Diameter 1.3 mm. However, transferring the authorship of this species makes Trochammina elegans Grzybowski, 1898 a primary junior homonym of Trochammina elegans Egger, 1893 (Article 53c of the ICZN). As such, the name must be permanently rejected (Article 52a). Because no junior synonym exists, we propose the replacement name Trochamminoides grzybowskii Kaminski & Geroch, nom. nov. SYSTEMATIC DESCRIPTION Suborder:Textulariina Delage & Herouard, 1896 Genus: Trochamminaides Cushman, 1910 Trochamminoides grzybowskii Kaminski & Geroch, nom.nov. for Trochammina elegans. . .


I N T R O D U C T I O N
Trochamminoides elegans (Grzybowski) [ = Trochammina elegans Grzybowski, 18981 is a familiar species that is commonly recorded from the Paleogene of the Carpathian flysch. However, during the course of work on a taxonomic revision of the Grzybowski Collection, it has become apparent that the name of this taxon is in actuality, invalid .
Thedesignation Trochamminaelegans was first used by Rzehak (1887), but since the name only appeared in a table (without a description or illustration) the name is unavailable under Article 12 of the ICZN. Grzybowski (1898), who had received specimens from Rzehak, referred to this species as "Trochamminaeleguns Rzehak,but theauthorship of the species belongs to Grzybowski under Article 50 of the ICZN. Grzybowski (1898) described Trochammina elegans as follows: Test agglutinated, surface rough; circular in outline, spirally coiled intwoand one-half to three whorls. Chambers spherical, increasing in size, numerous, with depressed sutures; there are thirteen to fifteen of them in the last whorl. Aperture at the lower edge of the last chamber, just above the rounded periphery. Diameter 1.3 mm.
However, transferring the authorship of this species makes Trochammina elegans Grzybowski, 1898 a primary junior homonym of Trochammina elegans Egger, 1893 (Article 53c of the ICZN). As such, the name must be permanently rejected (Article 52a). Because no junior synonym exists, we propose the replacement name Trochamrninoides grzybowskii Kaminski & Geroch, nom. nov.

SYSTEMATIC DESCRIPTION
Genus: Trochamminaides Cushman, 1910 Trocharnminoides grzybowskii Kaminski & Geroch, nom.nov. for Trochammina eleguns Grzybowski, 1898 (Fig 1) Derivation of Name. in honour of Jozef Grzybowski, who first described this taxon as a valid species. Type Specimen. The Grzybowski collection is housed in the micropaleontological collections of the Jagiellonian University in Krakow, Poland. The specimen of Trochammina elegans figured by Grzybowski is not preserved in the collection, but a n unfigured syntype is registered as 1 /58. We designate this specimen as the lectotype of Trochammina elegans Grzybowski, and hence under Article 72e it becomes the holotype of Trochamrninoides grzybowskii. Type Level. Eocene, Silesian unit of the Polish Outer Carpathians. The holotype is from gray clay collected along the bank of the Wislok River in Kroscienko Nizne, near Krosno, Poland. Description. Test large, oval in outline, with many chambers.

STANISLAW GEROCH
&&jute of Geological Sciences, lagiellonian University, ul. Oleandry 2a, 30-063 Krakoui, Poland Coiling is slightly glomospirai to planispiral, with approximately three whorls. Chambers are spherical, tightly appressed, numerous, increase in size slowly, about 15 in the last whorl. In immersion, the spherical chamber interiors are seen to beconnected by a thin tubedenoting anareal aperture. Wall thick, medium to finely agglutinated. Aperture small, just above the base of the last chamber. Dimensions. Diameter of holotype: 0.85 mm. Remarks. This species displays variability in the nature of coiling and in the number of chambers that is typical of the genus Trochamminoides. Grzybowski's figure depicts a planispirally coiled, laterally compressed specimen. However, the specimen preserved in his collection has distinct glomospiral coiling [i.e. coiled about a wobbly plane] in the early part of the test. It also has a more broadly rounded periphery than the type figure, is more oval in outline, and has a damaged final chamber. This specimen also departs from those illustrated by subsequent authors, who show round planispiral specimens. Our specimens from Zumaya are coiled planispirally and have more inflated chambers than the holotype. Rare specimens display a tendency to uncoil.