Abstract. Everticyclammina kelleri (Henson) (= E. eccentrica Redmond and E. elegans Redmond), a Berriasian-Valanginian index in the Middle East, is descended from the Late Jurassic Tethyan E. praekelleri sp.nov, a species which probably had its ancestry in Late Jurassic Ammobaculites sp. From this ancestor also evolved E. virguliana, and, in the Middle East, its Early Cretaceous descendants E. hensoni Redmond, E. contorta Redmond and E. greigi (Henson), which themselves formed the ancestry of the Albian Hemicyclammina whitei (Henson) and the Tethyan Albian - Cenomanian index H. sigali (Maync). Another, independent lineage from Ammobaculites, Buccicrenata hedbergi (Maync) (= B. libyca Gohrbandt) to B. subgoodlandensis (Vanderpool), transglobal in the Barremian - Cenomanian, is distinguished.